“The study covered some 850 swamp sites of about 110,000 hectares, including the Kushiro wetlands on Hokkaido and the Oze marshland, which stretches over Fukushima, Gunma and Niigata prefectures on Honshu.
It also covered around 1,300 sites classified as mud flats, totaling 50,000 hectares and including the Fujimae tidal flat in Aichi Prefecture on Honshu and the Ariake Sea, bordered by Fukuoka, Saga, Nagasaki and Kumamoto prefectures on Kyushu.”